Saints announced an agreement in principle with the French club last week, and the player’s medical and relevant paperwork have since been completed.
Mara said: “I’m really happy to sign for Southampton and to be a Saint for the next years. Southampton have given me a good opportunity to progress.
“I’m a young player and I have to work. I think Southampton is the best club for me to develop myself and progress to be one of the best. I’m impatient to begin with this team and score goals.”
Southampton manager Ralph Hasenhuttl added: “Sekou is coming here to score goals and has shown in the French league he can do this, now he needs to work hard and show he can do this in the Premier League.
“He is a young guy who will need time to adapt to a new country and a new league, but we are excited about his potential and what he can become in the future.”
The France U21 international becomes Saints' sixth summer signing following the arrivals of goalkeepers Gavin Bazunu and Mateusz Lis, defender Armel Bella-Kotchap, and midfielders Romeo Lavia and Joe Aribo.
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